Please email me if you would like a reprint:

Refereed Publications:


22. Riters, L.V., Spool, J.A., Merullo, D.P. & Hahn, A.H. (in press 5 Oct 2017). Song practice as a rewarding form of play in songbirds. Behavioural Processes [Invited].

21. Scully, E.N., Hahn, A.H., Campbell, K.A., McMillan, N., Congdon, J.V., & Sturdy, C.B. (in press) ZENK expression following conspecific and heterospecific playback in the zebra finch auditory forebrain. Behavioural Brain Research

20. Hahn, A.H., Campbell, K.A., Congdon, J.V., Hoang, J., McMillan, N., Scully, E.N., Yong, J.J.H., Elie, J.E., & Sturdy, C.B. (in press). Discrimination of acoustically similar conspecific and heterospecific vocalizations by black-capped chickadees (Poecile atricapillus). Animal Cognition. doi:10.1007/s10071-017-1087-5

19. McMillan, N., Hahn, A.H., Congdon, J.V., Campbell, K.A., Hoang, J., Scully, E.N., Spetch, M.L., & Sturdy, C.B. (in press). Chickadees discriminate contingency reversals presented consistently, but not frequently. Animal Cognition. doi: 10.1007/s10071-017-1088-4

18. Mercado, E. III, Wisniewski, M.G., McIntosh, B., Guillette, L.M., Hahn, A.H., & Sturdy, C.B. (in press). Chickadee songs provide hidden cues to singers’ locations. Animal Behavior and Cognition.

17. Hahn, A.H., Guillette, L.M., Hoeschele, M., Otter, K.A., Ratcliffe, L.M., & Sturdy, C.B. (2017). Discrimination of male black-capped chickadee songs: Relationship between acoustic preference and performance accuracy. Animal Behaviour, 126, 107-121. doi:10.1016/j.anbehav.2017.02.001

16. Hahn, A.H., Merullo, D.P., Spool, J.A., Angyal, C.S., Stevenson, S.A., & Riters, L.V. (2017). Song-associated reward correlates with endocannabinoid-related gene expression in male European starlings (Sturnus vulgaris). Neuroscience, 346, 255-266. doi:10.1016/j.neuroscience.2017.01.028

15. Sturdy, C. B., Campbell, K. A., Congdon, J. V., Hahn, A. H., McMillan, N., & Scully, E. N. (2017). Moving from perceptual to functional categories in songbirds. International Journal of Comparative Psychology. doi:10.7939/R3XP6VF7Z


14. Congdon, J.V., Hahn, A.H., McMillan, N., Avey, M.T., & Sturdy, C.B. (2016). Chickadee behavioural response to varying threat levels of predator and conspecific calls. Special Issue for the International Journal of Comparative Psychology.

13. Campbell, K.A., Hahn, A.H., Congdon, J.V., & Sturdy, C.B. (2016). Sex differences in black-capped chickadee (Poecile atricapillus) chick-a-dee calls. The Journal of the Acoustical Society of America. 140, 1598-1608. doi:10.1121/1.4962281

12. Hahn, A.H., Hoeschele, M., Guillette, L.M., Hoang, J., McMillan, N., Congdon, J.V., Campbell, K.A., Mennill, D.J., Otter, K.A., Grava, T., Ratcliffe, L.M., & Sturdy, C.B. (2016). Black-capped chickadees categorize songs based on features that vary geographically. Animal Behaviour, 112, 93-104. doi:10.1016/j.anbehav.2015.11.017


11. Hahn, A.H., Guillette, L.M., Lee, D., McMillan, N., Hoang, J., & Sturdy, C.B. (2015). Experience affects immediate early gene expression in response to conspecific call notes in black-capped chickadees (Poecile atricapillus). Behavioural Brain Research, 287, 49-58. doi:10.1016/j.bbr.2015.03.021

10. Hahn, A.H., Hoang, J., McMillan, N.A., Campbell, K., Congdon, J., & Sturdy, C.B. (2015). Biological salience influences performance and acoustic mechanisms for the discrimination of male and female songs. Animal Behaviour, 104, 213-228.  doi:10.1016/j.anbehav.2015.03.023

9. McMillan, N., Hahn, A.H., Spetch, M.L., & Sturdy, C.B. (2015). Avian cognition: Examples of sophisticated capabilities in space and song. WIRES Cognitive Science. doi: 10.1002/wcs.1346

8. Guillette, L.M., Hahn, A.H., Hoeschele, M., Przyslupski, A.-M., & Sturdy, C.B. (2015). Evidence for speed-accuracy trade-offs in cognition and personality in black-capped chickadees. Animal Cognition, 18, 165-178. doi: 10.1007/s10071-0140787-3


7. Zhai, Y., Dasog, M., Snitynsky, R.B., Purkait, T.K., Aghajamali, M., Hahn, A.H., Sturdy, C.B., Lowary, T.L., & Veinot, J.G.C. (2014). Water-soluble photoluminescent D-mannose and L-alanine functionalized silicon nanocrystals and their application to cancer cell imaging. Journal of Materials Chemistry B, 2, 8427-8433. doi: 10.1039/C4TB01161A

6. Hoeschele, M., Cook, R.G., Guillette, L.M., Hahn, A.H., & Sturdy, C.B. (2014). Timbre influences chord discrimination in black-capped chickadees but not humans. Journal of Comparative Psychology. doi: 10.1037/a0037159


5. Hahn, A.H., Guillette, L.M., Hoeschele, M., Mennill, D.J., Otter, K.A., Grava, T., Ratcliffe, L.M., & Sturdy, C.B. (2013). Dominance and geographic information contained within black-capped chickadee (Poecile atricapillus) song.  Behaviour, 150, 1601-1622. doi: 10.1163/1568539X-00003111

4. Hahn, A.H., Krysler, A., & Sturdy, C.B. (2013). Female song in black-capped chickadees (Poecile atricapillus): Acoustic song features that contain individual identity information and sex differences. Behavioural Processes, 98, 98-105. doi: 10.1016/j.bbr.2011.03.031

3. Hoeschele, M., Weisman, R., Guillette, L.M., Hahn, A.H., & Sturdy, C.B. (2013). Chickadees fail standardized operant tests for octave equivalence. Animal Cognition, doi: 10.1007/s10071-013-0597-z

2. Guillette, L.M., Hoeschele, M., Hahn, A.H., & Sturdy, C.B. (2013). Heterospecific discrimination of Poecile vocalizations by zebra finches (Taeniopygia guttata). Journal of Comparative Psychology, 127 (3), 227-236.  doi: 10.1037/a0029992


1. Hoeschele, M., Cook, R.G., Guillette, L.M., Hahn, A.H., & Sturdy, C.B. (2012). Auditory same/different concept learning and generalization in black-capped chickadees (Poecile atricapillus). PLoS ONE, 7(10), 1-11. doi:10.1371/journal.pone.0047691

Refereed Book Chapters:


2. Hahn, A.H., Congdon, J.V., Campbell, K.A., Scully, E.N., McMillan, N., & Sturdy, C.B. (2017). Mechanisms of Communication and Cognition in Chickadees: Explaining Nature in the Lab and Field. In M. Naguib, J. Podos, L. W. Simmons, L. Barrett, S. Healy, & M. Zuk (Eds.), Advances in the Study of Behavior (pp. 147–197).

1. McMillan, N., Avey, M.T., Bloomfield, L.L., Guillette, L.M., Hahn, A.H., Hoeschele, M., & Sturdy, C.B. (in press). Avian Vocal Perception: Bioacoustics and Perceptual Mechanisms. In C. ten Cate & S. D. Healy (Eds) Avian cognition. Cambridge University Press.

Non-refereed Publications:


2. Hahn, A.H., & Sturdy, C.B. (2015). Seemingly simple songs: Black-capped chickadee song revisited. Lay Language Paper for the Meeting of the Acoustical Society of America. Available online at:


1. Hahn, A.H., Guillette, L.M., Hoeschele, M., Cook, R.G., & Sturdy, C.B. (2013). Categories, concepts, and calls: Auditory perceptual mechanisms and cognitive abilities across different types of birds. Proceedings of Meetings on Acoustics, 19, doi: 10.1121/1.4799329

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